Phoenix Group

Evolutionary Ecology & Genetics

@ Te Taha Tawhiti Massey

Quenu M, Judd EJ, Morgan-Richards M, Trewick SA, Holt K, Tyler J, Morrey AM. 2023. High‐resolution stable isotope profiles from shells of the land snail Placostylus reveal contrasting patterns between snails originating from New Zealand and New Caledonia. Journal of Quaternary Science PDF
Meza-Joya FL, Morgan-Richards M, Koot EM, Trewick SA. 2023. Global warming leads to habitat loss and genetic erosion of alpine biodiversity. Journal of Biogeography 50: 961–975 PDF
Yarita S, Morgan-Richards M, Trewick SA. 2023. Genotypic detection of barriers to rat dispersal: Rattus rattus behind a peninsula predator proof fence. Biological Invasions PDF
Morgan-Richards M, Marshall C, Biggs PJ, Trewick SA. 2023. Insect Freeze-Tolerance Downunder: The Microbial Connection. Insects 14, 89 PDF
Carmelet-Rescan D, Morgan-Richards M, Pattabiraman N, Trewick SA. 2022. Time-calibrated phylogeny and ecological niche models indicate Pliocene aridification drove intraspecific diversification of brushtail possums in Australia. Ecology & Evolution 12: e9633 PDF
Trewick S, Hegg D, Morgan-Richards M, Murray T, Watts C, Johns P, Michel P. 2022. Conservation status of
Orthoptera (wētā, crickets and grasshoppers) in Aotearoa New Zealand. New Zealand Threat Classification
Series 39. Department of Conservation, Wellington. 28 p. NZTCS
Godfrey AJR, McKenzie AO, Morgan-Richards M. 2022. Recommendations for non-lethal monitoring of tree wētā (Hemideina spp.) using artificial galleries. New Zealand Journal of Zoology PDF
Nakano M, Morgan-Richards M, Clavijo-McCormick A, Trewick SA. 2022. Abundance and distribution of antennal sensilla on males and females of three sympatric species of alpine grasshopper (Orthoptera: Acrididae: Catantopinae) in Aotearoa New Zealand. Zoomorphology PDF
Cunha A, Carmelet-Rescan D, Marques AC, Morgan-Richards M. 2022. Contrasting morphological and genetic patterns suggest cryptic speciation and phenotype-environment covariation within three benthic marine hydrozoans. Marine Biology. 169: 103. PDF
Trewick SA, Hendersion IM, Pohe SR, Morgani-Richards M. 2022. Spatial Variation of Acanthophlebia cruentata (Ephemeroptera), a Mayfly Endemic to Te Ika-a-Māui—North Island of Aotearoa, New Zealand. Insects 13(7) 1755102  PDF
Meza-Joya FL, Morgan-Richards M, Trewick SA. 2022. Relationships among body size components of three flightless New Zealand grasshopper species (OrthopteraAcrididae) and their ecological applications. Journal of Orthoptera Research 21: 91–103 PDF
Bhavanam S, Trewick SA. 2022. Effects of population density on adult morphology and life-history traits of female Mediterranean flour moth, Ephestia kuehniella (Lepidoptera: Pyralidae). European Journal of Entomology 119: 191–200. PDF
Shepherd L, Simon C, Langton-Myers S, Morgan-Richards M. 2022. Insights into Aotearoa New Zealand’s biogeographic history provided by the study of natural hybrid zones. Journal of the Royal Society of New Zealand Online PDF
Hegg D, Morgan-Richards M, Trewick SA. 2022. High alpine sorcerers: revision of the cave wētā genus Pharmacus
Pictet & de Saussure (Orthoptera: Rhaphidophoridae: Macropathinae), with the description of six new species and three new subspecies. European Journal of Taxonomy 808: 1–58
Koot EM, Morgan-Richards M, Trewick SA. 2022. Climate change and alpine-adapted insects: modelling environmental envelopes of a grasshopper radiation. Open Science 9: 211596 PDF
Nakano M, Morgan-Richards M, Trewick SA, Clavijo McCormick A. 2022. Chemical ecology and olfactory in short-horned grasshoppers (Orthoptera: Acrididae). Journal of Chemical Ecology 48: 121–140 PDF
Pattabiraman N, Morgan-Richards M, Powlesland R, Trewick SA. 2021. Unrestricted gene flow between two subspecies of translocated brushtail possums (Trichosurus vulpecula) in Aotearoa New Zealand. Biological Invasions 24: 247–260 PDF
Quenu M, Trewick SA, Daly EE, Morgan-Richards M. 2021. Generation of large mitochondrial and nuclear nucleotide sequences and phylogenetic analyses using high-throughput short-read datasets for endangered Placostylinae snails of the southwest Pacific. Molluscan Research 41: 243–253 PDF
Morgan-Richards M, Vilcot M, Trewick SA. 2021 Lack of assortative mating might explain reduced phenotypic differentiation where two grasshopper species meet. Journal of Evolutionary Biology 35: 509–519 PDF
Stasiak K, Dunowska M, Trewick S, Rola J. 2021. Genetic variation in the glycoprotein B sequence of equid herpesvirus 5 among horses of various breeds at Polish national studs. Pathogens 10: 322 PDF
Trewick SA. 2021. A new species of large Hemiandrus ground wētā (Orthoptera: Anostostomatidae) from North Island,
New Zealand. Zootaxa 4942 (2)
Carmelet-Rescan D, Morgan-Richards M, Koot EM, Trewick SA. 2021. Climate and ice in the last glacial maximum explain patterns of isolation by distance inferred for alpine grasshoppers. Insect Conservation & Diversity 14: 568–581 PDF
Wehi PM, Brownstein G, Morgan-Richards M. 2020. Indigenous plant naming and experimentation reveal a plant–insect relationship in New Zealand forests. Conservation Science in Practice 2: e282 PDF
Trewick SA, Taylor-Smith BL, Morgan-Richards M. 2020. Ecology and systematics of the wine wētā and allied species, with description of four new Hemiandrus species. New Zealand Journal of Zoology 48: 47–80 PDF
Morgan-Richards M, Trewick SA, Cox M. 2020. Recent evidence in human evolution. NZ Association of Science Educators. PDF
Daly EE, Trewick SA, Dowle EJ, Crampton JS, Morgan-Richards M. 2020. Conservation of pūpū whakarongotaua - the snail that listens for the war party. Ethnobiology & Conservation 9: 13. PDF
Gaspar J, Gibb GC, Trewick SA. 2020. Convergent morphological responses to loss of flight in rails (Aves: Rallidae). Ecology & Evolution 10: 6186–6207  PDF
Vaux F, Gemmell MR, Hills SFK, Marshall BA, Beu AG, Crampton JS, Trewick SA, Morgan-Richards M. 2020. Lineage identification affects estimates of evolutionary mode in marine snails. Systematic Biology 69: 1106–1121. PDF
Koot EM, Morgan-Richards, M, Trewick SA. 2020. An alpine grasshopper radiation older than the mountains, on Kā Tiritiri o te Moana (Southern Alps) of Aotearoa (New Zealand). Molecular Phylogenetics & Evolution 147: 106783 PDF
Taylor-Smith BL, Morgan-Richards M, Trewick SA. 2019. Patterns of regional endemism among New Zealand invertebrates. New Zealand Journal of Zoology 47: 1–19 PDF
Quenu M, Trewick SA, Brescia F, Morgan-Richards M. 2020. Geometric morphometrics and machine learning challenges the division of Placostylus snails from the Isle of Pines, New Caledonia into just two species. Journal of Molluscan Studies 86: 35–41. PDF
Nakano M, Morgan-Richards M, Godfrey AJR, Clavijo McCormick A. 2019. Parthenogenetic females of the stick insect Clitarchus hookeri maintain sexual traits. Insects 10: 202 
Hegg D, Morgan-Richards M, Trewick SA. 2019. Diversity and distribution of Pleioplectron Hutton cave wētā (Orthoptera: Rhaphidophoridae: Macropathinae), with synonymy of Weta Chopard and description of seven new species. European Journal of Taxonomy #577 PDF
Gemmell M, Trewick SA, Hills SFK, Morgan-Richards M. 2019. Phylogenetic topology and timing of the New Zealand olive shells is consistent with Punctuated Equilibrium. Journal of Zoological Systematics and Evolutionary Research 58: 209–220 PDF
Morgan-Richards M, Langton-Myers S, Trewick SA. 2019. Loss and gain of sexual reproduction in the same stick insect. Molecular Ecology 28: 3929–3941.  PDF
Daly EE, Walker KJ, Morgan-Richards M, Trewick SA. 2019. Spatial genetics of a high elevation lineage of Rhytididae land snails in New Zealand: the Powelliphanta Kawatiri complex. Molluscan Research 39: 280–289. PDF
Vaux F, Morgan-Richards M, Daly EE, Trewick SA. 2018. Tuatara and a new morphometric dataset for Rhynchocephalia: comments on Herrera-Flores et al. Palaeontology 62: 321–334. PDF
Gemmell M, Trewick SA, Crampton JS, Vaux F, Hills SFK, Daly EE, Marshall BA, Beu AG, Morgan-Richards M. 2018. Genetic structure and shell shape variation within a rocky shore whelk suggest both diverging and constraining selection with gene flow. Biological Journal of the Linnean Society 125: 827–843. PDF
Bulgarella M, Quenu M, Shepherd LD, Morgan-Richards M. 2018 The ectoparasites of hybrid ducks in New Zealand (Mallard x Grey Duck). Parasites & Wildlife 7: 335–342. PDF
Fitness J, Morgan-Richards M, Hegg D, Trewick SA. 2018. Reinstatement of the New Zealand cave wētā genus Miotopus Hutton (Orthoptera: Rhaphidophoridae) and description of a new species. European Journal of Taxonomy #468. PDF
Garcia‐R JC, Gonzale Orosco, CE, Trewick SA. 2019. Contrasting patterns of diversification in a bird family (Aves: Gruiformes: Rallidae) are revealed by analysis of geospatial distribution of species and phylogenetic diversity. Ecography 42: 500–510. PDF
McKean NE, Trewick SA, Griffin MJ, Dowle EJ, Morgan-Richards M. 2018. Viability and fertility of hybrid New Zealand tree wētā Hemideina spp. (Orthoptera: Anostostomatidae). Journal of Orthoptera Research 27(2): 97–106 PDF
Vaux F, Trewick SA, Crampton JS, Marshall BA, Beu AG, Hills SFK, Morgan-Ricahrds M. 2018. Evolutionary lineages of marine snails identified using molecular phylogenetics and geometric morphometric analysis of shells. Molecular Phylogenetics & Evolution 127: 626–637. PDF
Bridgeman B, Morgan-Richards M, Wheeler D, Trewick SA. 2018. First detection of Wolbachia in the New Zealand biota. PLoS ONE 13 (4): e0195517. PDF
Alfaro FM, Muñoz-Ramírez CP, Zúñiga-Reinoso A, Trewick SA, Méndez MA. 2018. Phylogeography of the Chilean red cricket Cratomelus armatus (Orthoptera: Anostostomatidae) reveals high cryptic diversity in central Chile. Biological Journal of the Linean Society 123: 712–727 PDF
Trewick SA, Morgan-Richards M. 2018. Missing New Zealand stickman found in UK. Antenna 42: 10–13.
(Reproduced with permission of Antenna, the bulletin of the Royal Entomological Society)
Sivyer L, Morgan-Richards M, Koot E, Trewick SA. 2018. Anthropogenic cause of range shifts and gene flow between two grasshopper species revealed by environmental modelling, geometric morphometrics and population genetics. Insect Conservation & Diversity 11: 415–434. PDF
Marshall BA, Hills SFK, Vaux F. 2018. A new species of Penion P. Fischer, 1884 from northern New Zealand (Mollusca: Neogastropoda: Buccinoidea). Molluscan Research 38: 238–242. PDF
Vaux F, Hills SFK, Marshall BA, Trewick SA, Morgan-Richards M. 2018. Genome statistics and phylogenetic reconstructions for Southern Hemisphere whelks (Gastropoda: Buccinulidae). Data in Brief 16: 172–181. PDF
Brock PD, Lee M, Morgan-Richards M, Trewick SA. 2018. Missing stickman found: the first male of the parthenogenetic New Zealand Phasmid genus Acanthoxyla Uvarov, 1944 discoverd in the United Kingdom. Atropos 60: 16–23. PDF
Morgan-Richards M, Bulgarella M, Sivyer L, Dowle E, Hale M, McKean N, Trewick SA. 2017. Explaining large mitochondrial sequence differences within a population sample. Royal Society Open Access 4(11), 170730. PDF
Trewick SA, Pilkington S, Shepherd LD, Gibb GC, Morgan-Richards M. 2017. Closing the gap: avian lineage splits at a young, narrow seaway imply a protracted history of mixed population response. Molecular Ecology 26: 5752–5772. PDF
Wehi PM, Monks A, Morgan-Richards M. 2017. Male tree weta are attracted to cuticular scent cues but do not discriminate according to sex or among two closely related species. Ethology 123: 825–834. PDF
Trewick SA. 2017 Plate tectonics in biogeography. In, The International Encyclopedia of Geography (Eds. Richardson D. et al.). John Wiley & Sons Ltd. PDF
Vaux F, Crampton JS, Marshall BA, Trewick SA, Morgan-Richards M. 2017. Geometric morphometric analysis reveals that the shells of male and female siphon whelks Penion chathamensis are the same size and shape. Molluscan Research 37: 194–201. PDF
Garcia-R JC, Joseph L, Adcock G, Reid J, Trewick SA. 2017. Interisland gene flow among populations of the buff-banded rail (Aves: Rallidae) and its implications for insular endemism in Oceania. Journal of Avian Biology 48: 679–690. PDF
Vaux F, Hills SFK, Marshall BA, Trewick SA, Morgan-Richards M. 2017 A phylogeny of Southern Hemisphere whelks (Gastropoda: Buccinulidae) and concordance with the fossil record. Molecular Phylogenetics & Evolution 114: 367–381. PDF
Bhavanam S, Trewick SA. 2017. Effects of larval crowding and nutrient limitation on male phenotype, reproductive investment and strategy in Ephestia kuehniella Zeller (Insecta: Lepidoptera). Journal of Stored Products Research 71: 64–71. PDF
Bradley E, Trewick SA, Morgan-Richards M. 2017. Genetic distinctiveness of the Waikawa Island mice population indicates low rate of dispersal from mainland New Zealand. New Zealand Journal of Ecology 41: 245–250. PDF
Trewick SA, Olley L. 2016. Spatial size dimorphism in New Zealand’s last endemic raptor, the Kārearea Falco novaeseelandiae, coincides with a narrow sea strait. IBIS 158: 747–761. PDF
Trewick SA, Johns P, Hitchmough R, Rolfe J, Stringer I. 2016. Conservation status of New Zealand Orthoptera, 2014. New Zealand Threat Classification Series 16. Department of Conservation, Wellington. 15 p. PDF
Vaux F., Trewick S.A, Morgan-Richards M. 2016. Speciation through the looking-glass. Biological Journal of the Linnean Society 120: 480–488. PDF
Taylor-Smith BL, Trewick SA, Morgan-Richards M. 2016. Three new ground wētā and a redescription of Hemiandrus maculifrons. New Zealand of Zoology 43: 363–383. PDF
Yabaki M, Winkworth RC, McLenachan PA, Aalbersberg W, Winder L, Trewick SA, Lockhart PJ. 2016. Placing the Fijian Honeyeaters within the meliphagid radiation: implications for origins and conservation. Pacific Conservation Biology 22: 262–271. PDF
Doddala PRC, Minor MA, Wang Q, Rogers DJ, Koot EM, Trewick SA. 2016. Role of olfaction in host plant selection and local adaptation of a polyphagous herbivore, Eucolaspis Sharp. J. Applied Entomology 140: 444–452. PDF
Morgan-Richards M, Hills, SFK, Biggs PJ, Trewick SA. 2016. Sticky genomes: using NGS evidence to test hybrid speciation hypothesis. PLOS One 11(5): e0154911. PDF
McKean NE, Trewick SA, Morgan-Richards M. 2016. Little or no gene flow despite F1 hybrids at two interspecific contact zones. Ecology & Evolution 6: 2390–2404. PDF
Morgan-Richards M, Hinlo AR, Smuts-Kennedy C, Innes J, Ji W, Barry M, Brunton D, Hitchmough RA. 2016. Identification of a rare gecko from North Island New Zealand, and genetic assessment of its probable origin: A novel mainland conservation priority? Journal of Herpetology 50: 77–86. PDF
Nelson-Tunley M, Morgan-Richards M, Trewick SA. 2016. Genetic diversity and gene flow in a rare New Zealand skink despite fragmented habitat in a volcanic landscape. Journal of the Linnean Society 119: 37–51. PDF
Trewick SA, Morgan-Ricahards M. 2016. Phylogenetics and Conservation in New Zealand: The Long and the Short of It. Pp 81–97, in Biodiversity Conservation and Phylogenetic Systematics (eds. Pellens, R., Grandcolas P.). Topics in Biodiversity & Conservation 14. Springer. Online PDF
Grandcolas P, Trewick SA 2016. What Is the Meaning of Extreme Phylogenetic Diversity? The Case of Phylogenetic Relict Species. Pp 99–115, in Biodiversity Conservation and Phylogenetic Systematics (eds. Pellens, R., Grandcolas P.). Topics in Biodiversity & Conservation 14. Springer. Online  
Garcia‐R JC, Elliott G, Walker K, Castro I, Trewick SA. 2016. Trans-equatorial range of a land bird lineage (Aves: Rallidae) from tropical forests to subantarctic grasslands. Journal of Avian Biology 47: 219–226. PDF
Doddala PRC, Minor MA, Rogers DJ, Trewick SA. 2015. Fifteen into three does go: Morphology, genetics and genitalia confirm taxonomic inflation of New Zealand beetles (Chrysomelidae: Eucolaspis). PLoS ONE 10(11): e0143258. PDF
Sangster G, García-R JC, Trewick SA. 2015. A new genus for the Lesser Moorhen Gallinula angulata Sundevall, 1850 (Aves, Rallidae). European Journal of Taxonomy 153: 1–8. PDF
Gibb GC, England R, Hartig G, McLenachan PA, Taylor Smith BL, McComish BJ, Cooper A, Penny D. 2015. New Zealand passerines help clarify the diversification of major songbird lineages during the Oligocene. Genome Biology and Evolution 7: 2983–2995. PDF
Doddala PRC, Minor MA, Wang Q, Koot EM, Trewick SA. 2015. Role of olfaction in host plant selection and local adaptation of a polyphagous herbivore, Eucolaspis Sharp. Journal of Applied Entomology 140: 444–452. PDF
Bulgarella M, Trewick SA, Godfrey AJR, Sinclair BJ, Morgan-Richards M. 2015. Elevational variation in adult body size and growth rate but not in metabolic rate in the tree weta Hemideina crassidens. Journal of Insect Physiology 75:30–38. PDF
Gibb GC, Condamine FL, Kuch M, Enk J, Moraes-Barros N, Superina M, Poinar HN, Delsuc F. 2015. Shotgun mitogenomics provides a reference phylogenetic framework and timescale for living xenarthrans. Molecular Biology & Evolution 33: 621–642. PDF
Steel D, Stewart BG, Trewick SA, Wallis GP. 2015. Fiord populations of Astrobrachion constrictum (Ophiuroidea: Asteroschematidae) show little genetic differentiation for mitochondrial DNA. New Zealand Journal Zoology 42: 165–172. PDF
Vaux F, Trewick SA, Morgan-Richards M. 2015. Lineages, splits and divergence challenge whether the terms anagenesis and cladogenesis are necessary. Biological Journal of the Linnean Society 117: 165–176. PDF
Dowle EJ, Morgan-Richards M, Brescia F, Trewick SA. 2015. Correlation of shell phenotype and local environment suggests a role for natural selection in the evolution of Placostylus snails. Molecular Ecology 24: 4205–4221. PDF
Grandcolas P, Nattier R, Trewick SA. 2014. Relict species: a relict concept? Trends in Ecology & Evolution 29: 655–663. PDF
Goldberg J, Morgan-Richards M, Trewick SA. 2015. Intercontinental island hopping: colonization and speciation of the grasshopper genus Phaulacridium (Orthoptera: Acrididae) in Australasia. Zoologisher Anzeiger 255: 71–79. PDF
McKean N, Morgan-Richards M, Trewick SA. 2015. Comparative cytogenetics of North Island tree weta in sympatry. New Zealand Journal of Zoology 42: 73–84. PDF
Fitness J, Morgan-Richards M, Ball O, Godfrey J, Trewick SA. 2015. Improved resolution of cave weta diversity (Orthoptera: Rhaphidophoridae): ecological implications for Te Paki, Far North, New Zealand. New Zealand Journal of Zoology 42: 1–16. PDF
Dowle EJ, Morgan-Richards M, Trewick SA. 2014. Morphological differentiation despite gene flow in an endangered grasshopper. BMC Evolutionary Biology 14: 216. PDF
García-R JC, Trewick SA. 2014. Dispersal and speciation in purple swamphens (Rallidae: Porphyrio). The Auk 132: 140–155 PDF
Mahmood MT, McLenachan PA, Gibb GC, Penny D. 2014. Phylogenetic position of avian nocturnal and diurnal raptors. Genome Biology and Evolution6: 326–332.  
García-R JC, Gibb G, Trewick SA. 2014. Eocene diversification of crown group rails (Aves: Gruiformes: Rallidae). PLoS ONE9:e109635. PDF
García-R JC, Gibb G, Trewick SA. 2014. Deep global evolutionary radiation in birds: diversification and trait evolution in the cosmopolitan bird family Rallidae. Molecular Phylogenetics & Evolution 81: 96–108. PDF
Trewick SA, Morgan-Richards M. 2014. New Zealand Wild Life. Penguin, New Zealand. ISBN-13:9780143568896 ISBN-10:0143568892 New edition  
Minards NZ, Trewick SA, Godfrey JR, Morgan-Richards M. 2014. Convergent local adaption in size and growth rate but not metabolic rate in a pair of parapatric Ortophtera species. Biological Journal of the Linnean Society 113: 123–135 PDF
Hinlo RP, Tabora JAG, Bailey CA, Trewick SA, Robeng G, van Weerd M, Pomares CC, Engberg SE, Brenneman RA, Louis EE. 2014. Population genetics implications for the conservation of the Philippine crocodile Crocodylus mindorensis Schmidt, 1935. Journal of Threatened Taxa 6: 5513–5533. PDF
Goldberg J, Knapp M, Emberson RM, Townsend JI, Trewick SA. 2014. Species radiation of Carabid beetles (Broscini: Mecodema) in New Zealand. PLoS ONE Online PDF
Bulgarella M, Trewick S, Minards NA, Jacobson MJ, Morgan-Richards M. 2014. Shifting ranges of two tree weta species (Hemideina spp.): competitive exclusion and changing climate. Journal of Bioleography41: 524–535. PDF
Gibb GC, Hills SFK. 2013. Intergenerational mutation rate does not equal long-term evolutionary substitution rate. Proceedings of the National Academy of Science USA. 110:e611  
Taylor Smith BL, Morgan-Richards M, Trewick SA. 2013. New Zealand ground wētā (Anostostomatidae: Hemiandrus): descriptions of two species with notes on their biology. New Zealand Journal of Zoology 40: 314–329. PDF
Doddala PRC, Trewick SA, Rogers DJ, Minor MA. 2013. Predictive modelling of adult emergence in a polyphagous Eucolaspis leaf beetle. Journal of Economic Entomology 106: 899–904. PDF
Gibb GC, Kennedy M, Penny D. 2013. Beyond phylogeny: Pelecaniform and Ciconiiform Birds, and long term niche stability. 68:229-238. Molecular Phylogenetics & Evolution68: 229–238. PDF
Wehi PM, Raubenheimer D, Morgan-Richards M. 2013. Tolerance for nutrient imbalance in an intermitently feeding herbivorous cricket, the Wellington tree weta. PLoS One 8:e84641 PDF
Dowle EJ, Morgan-Richards M, Trewick SA. 2013. Molecular evolution and the latitudinal biodiversity gradient. Heredity 110: 501–510. PDF
Waters JM, Trewick SA, Paterson AM, Spencer HG, Kennedy M, Craw D, Burridge CP, Wallis GP. 2013. Biogeography off the tracks Systematic Biology 62: 494–498. PDF
Kant R, Minor M, Sandanayaka M, Trewick SA. 2013. Effects of mating and oviposition delay on parasitism rate and sex allocation behaviour of Diaeretiella rapae (Hymenoptera: Aphidiidae). Biological Control 65: 265–270. PDF
Hills SFK, Crampton JS, Trewick SA, Morgan-Richards M. 2012. DNA and morphology unite two species and 10 million year fossils. PLoS One. 7(12): 1–13. PDF
Trewick SA, Bland K. 2012. Fire and slice: palaeogeography for biogeography at New Zealand’s North Island/South Island juncture. Journal of the Royal Society of New Zealand 42: 153–183. PDF
Myers S, Trewick SA, Morgan-Richards M. 2012. Multiple lines of evidence suggest mosaic polyploidy in the hybrid parthenogenetic stick insect lineage Acanthoxyla. Insect Conservation & Diversity. 6: 537–548. PDF
Tabora JAG, Hinlo RP, Bailey CA, and others. 2012. Detection of Crocodylus mindorensis x Crocodylus porosus (Crocodylidae) hybrids in a Philippine crocodile systematics analysis. Zootaxa 3560: 1–31.  
Kant R, Trewick SA, Sandanayaka WRM, Godfrey AJR, Minor MA. 2012. Effects of multiple matings on reproductive fitness of male and female Diaeretiella rapae. Entomologia Experimentalis et Applicata. 145: 215–221. PDF
Wehi PM, Jorgensen M. Morgan-Richards M. Sex and seasonal movement in a flightless forest insect the Auckland tree weta. New Zealand Journal of Ecology. 37: 75-83. PDF
Kant R, Minor MA and Trewick SA 2012. Mating or oviposition? A crucial decision in the life history of cabbage aphid parasitoid Diaeretiella rapae (McIntosh) (Hymenoptera: Aphidiidae) Ecological Entomology. 37: 169-174. PDF
Kant R, Minor MA and Trewick SA 2012. Reproductive strategies of Diaeretiella rapae (Hymenoptera: Aphidiinae) during fluctuating temperatures of spring season in New Zealand. Biocontrol Science & Technology. 22: 1-9. PDF
Wehi PM, Whaanga H, Trewick SA. 2012. Artefacts, biology and bias in museum collection research. Molecular Ecology 21: 3103-3109. Online PDF
Kant R, Minor MA, Trewick SA, Sandanayaka WRM. 2012. Body size and fitness relation in male and female Diaeretiella rapae. Biocontrol. 57(6):759-766 DOI 10.1007/sl0526-012-09452-4 PDF
Fitness J, Hitchmough RA, Morgan-Richards M. 2011. Little and large: body size and genetic clines in a New Zealand gecko (Woodworthia maculata) along a coastal transect. Ecology and Evolution. 2: 273-285. Online PDF
Kant R, Minor MA, Trewick SA. 2012 Fitness gain in a koinobiont parasitoid Diaeretiella rapae (Hymenoptera: Aphidiidae) by parasitising hosts of different ages. Journal of Asia-Pacific Entomology. 15: 83–87 PDF
Trewick SA. 2011. Vicars and vagrants: Assembly of the New Zealand avifauna. Australasian Science 32: 24-27. PDF
Hills SFK, Trewick SA, Morgan-Richards M. 2011 Phylogenetic information of genes, illustrated with mitochondrial data from a genus of gastropod molluscs. Biological Journal of the Linnean Society.104: 770-785. Online PDF
Chappell EM, Trewick SA, Morgan-Richards M. 2012 Shape and sound reveal genetic cohesion not speciation in the New Zealand orthopteran, Hemiandrus pallitarsis, despite high mtDNA divergence. Biological Journal of the Linnean Society. 105: 169-186. Online PDF
Wehi PM, Nakagawa S, Trewick SA, Morgan-Richards M. 2011. Does predation result in adult sex ratio skew in a sexually dimorphic insect genus? Journal of Evolutionary Biology. 24: 2321–2328. Online
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Goldberg J, Trewick SA. 2011 Exploring phylogeographic congruence in a continental island system. Insects.2: 369-399. Online. PDF
Kant R, Minor MA, Trewick SA, Sandanayaka WRM. 2011. Host selection for self-superparasitism  by Diaeretiella rapae (M’Intosh) (Hymenoptera: Aphidiidae). New Zealand Plant Protection 64: 37–43. Online PDF
Trewick SA, Wallis GP, Morgan-Richards M. 2011. The invertebrate life of New Zealand: a phylogeographic approach. Insects. 2: 297–325 Online PDF
Segovia R, Walker Pett W, Trewick SA, Lavrov DV. 2011. Extensive and evolutionarily persistent mitochondrial tRNA editing in velvet worms (phylum Onychophora). MBE. PDF
Griffin MJ, Morgan-Richards M, Trewick SA. 2011. Is the tree weta Hemideina crassidens an obligate herbivore? New Zealand Natural Sciences. 36: 11–19. PDF
Griffin MJ, Trewick SA, Wehi PM, Morgan-Richards M. 2011. Exploring the concept of niche convergence in a land without rodents: the case of weta as small mammals. New Zealand Journal of Ecology. 35(3): 302-307 PDF
Crisp M, Trewick SA, Cook LG. 2010. Hypothesis testing in biogeography. Trends in Ecology and Evolution 1317: 1–7. PDF
Goldberg J, Trewick SA, Powlesland RG. 2010. Population structure and biogeography of Hemiphaga pigeons (Aves: Columbidae) on islands in the New Zealand region. Journal of Biogeography 38: 285–298. PDF
Gibb GC, D Penny. 2010. Two aspects along the continuum of pigeon evolution: A South-Pacific radiation and the relationship of pigeons within Neoaves. Molecular Phylogenetics and Evolution 56: 698–706 Online  
Allegrucci G, Trewick SA, Forunato A, Carchini G, Sbordoni V. 2010. Cave crickets and cave weta (Orthoptera) from the southern end of the world: a molecular phylogeny test of biogeographical hypotheses. Journal of Orthoptera Research 19: 121–130. PDF
Wyman TE, Trewick SA, Morgan-Richards M, and Noble ADL. 2010. Mutualism or opportunism? Tree fuchsia (Fuchsia excorticata) and tree weta (Hemideina) interactions. Austral Ecology 36: 261–281. PDF
Cook LD, Morgan-Richards M, Trewick SA, Johns P. 2010. Status of the New Zealand cave weta (Rhaphidophoridae) genera Pachyrhamma, Gymnoplectron and Turbottoplectron. Invertebrate Systematics 24: 131–138. PDF
Morgan-Richards M, Trewick SA, Stringer IA. 2010. Geographic parthenogenesis and the common tea-tree stick insect of New Zealand. Molecular Ecology 19: 1227–1238. PDF
Trewick SA, Gibb G. 2010. Assembly of the New Zealand avifauna- a review of molecular evidence. IBIS 152: 226–253. PDF
Landis CA, Campbell HJ, Begg JG., Mildenhall DC, Paterson AM, Trewick SA. 2010. Discussion of ‘The Waipounamu Erosion Surface: questioning the antiquity of the New Zealand land surface and terrestrial. Geological  Magazine 147: 151–155. PDF
Macfarlane, R.P. et al. 2010 Phylum Arthropoda, subphylum Hexapoda. In: Gordon, D.P. (ed.) The New Zealand Inventory of Biodiversity. Volume 2. Kingdom Animalia: Chaetognatha, Ecdysozoa, Ichnofossils. Canterbury University Press, Christchurch.  
Doddala P.R.C., Minor M.A., and Rogers D.J. 2010. Population dynamics of bronze beetle, Eucolaspis spp (Coleoptera: Chrysomelidae) in relation to other soil macro-invertebrates in organic apple orchards in Hawke’s Bay, New Zealand. New Zealand Plant Protection 63:195–200 PDF
Morgan-Richards M, Smissen RD, Shepherd LD, Hayward J, Chan C, Chambers GK, Wallis GP, Chapman HM. 2009 A review of genetic analyses of hybridisation in New Zealand. Journal of the Royal Society of New Zealand 39: 15–34. PDF
Trewick SA, Morgan-Richards M. 2009 New Zealand Biology. Pages 665-673 in Encyclopedia of Islands (Eds R.G. Gillespie and D.A. Clague). University of California Press, Berkeley. PDF
Trewick SA. Te Ara- Encyclopedia of New Zealand, Ministry of Culture .  
Trewick SA, Brescia F, Jordan C. 2009. Diversity and phylogeny of New Caledonian Placostylus land snails; evidence from mitochondrial DNA. In Zoologia Neocaledonica 7. Biodiversity studies in New Caledonia. Mémoires du Muséum national d'Histoire naturelle, 196 (ed. P. Grandcolas), 295–310. PDF
Wallis GP, Trewick SA. 2009. New Zealand phylogeography: evolution on a small continent. Molecular Ecology, 18, 3548–3580. doi: 10.1111/j.1365-294X.2009.04294.x PDF
Pratt RC, Gibb GC, Morgan-Richards M, Phillips MJ, Hendy MD, Penny D. 2009. Towards resolving deep Neoaves phylogeny: data, signal enhancement and priors Molecular Biology and Evolution; 26(2): 313–326 PDF
Trewick SA, Cowie RH. 2008 Introduction. Evolution on Pacific islands: Darwin’s legacy. Philosophical Transactions of the Royal Society, London 363: 3289–3291. Go here for more information about content of this SPECIAL DARWIN200 EDITION PDF
Trewick SA, Morgan-Richards M, Collins LJ. 2008. Are you my mother? DNA reveals hybrid stick insect genus is an orphan. Molecular Phylogenetics and Evolution 48:799-808. PDF
Trewick SA, Walker KJ, Jordan CJ. 2008. Taxonomic and conservation status of a newly discovered giant landsnail from Mount Augustus, New Zealand. Conservation Genetics 9: 1563-1575. PDF
Walker KJ, Trewick SA, Barker GM. 2008. A new species of Powelliphanta (Mollusca: Pulmonata: Rhytididae) land snail from the Stockton Plateau, New Zealand. Journal of the Royal Society of New Zealand 38: 163–186. PDF
Trewick SA. 2008 DNA Barcoding is not enough: mismatch of taxonomy and genealogy in New Zealand grasshoppers (Orthoptera: Acrididae). Cladistics 24: 240-254. PDF
Gibb GC, Kardailsky O, Kimball RT, Braun EL, Penny D. 2007. Mitochondrial Genomes and Avian Phylogeny: Complex Characters and Resolvability without Explosive Radiations. Molecular Biology and Evolution 24:269-280. Online  
Trewick S, Morris S. 2008 Diversity and taxonomic status of some New Zealand grasshoppers. DOC Research & Development Series #290. Wellington, New Zealand. PDF
Pratt RC, Morgan-Richards M, Trewick SA. 2008. Diversification of New Zealand weta (Orthoptera: Ensifera: Anostostomatidae) and their relationships in Australasia. Philosophical Transactions Royal Society, London 363: 3427–3438. PDF
Trewick SA, Paterson AM, Campbell HJ. 2007. Hello New Zealand. Journal of Biogeography  34: 1–6. PDF
Goldberg J, Trewick SA, Paterson AM. 2008 Evolution of New Zealand's terrestrial fauna: a review of molecular evidence. Philosophical Transactions of the Royal Society, London 363: 3319–3334. PDF
Landis CA, Campbell HJ, Begg JG, Mildenhall DC, Paterson AM, Trewick SA. 2008 The Waipounamu Erosion Surface:  Questioning the Antiquity of the New Zealand Land Surface and Terrestrial Fauna and Flora. Geological Magazine 145: 1–25. PDF
Morgan-Richards MTrewick SA, Bartosch-Härlid A, Kardailsky O, Phillips MJ, McLenachan PA, Penny D. 2008. Bird Evolution: Testing The Metaves Clade With Six New Mitochondrial Genomes. BMC Evolutionary Biology 8:20. PDF
Morgan-Richards M, Trewick SA, Dunavan S. 2008. When is it coevolution? The case of ground wētā and fleshy fruits in New Zealand. New Zealand Journal of Ecology 32: 108–112. PDF
Neall VE, Trewick SA. 2008. The age and origin of the pacific islands - a geological overview. Philosophical Transactions of the Royal Society, London 363: 3293–3308. PDF
Morgan-Richards M. 2005. Chromosome rearrangements are not accompanied by expected genome size change in the tree weta Hemideina thoracica (Orthoptera, Anostostomatidae). Journal of Orthoptera Research 14(2): 143–148. PDF
Morgan-Richards M, Trewick SA. 2005. Hybrid origin of a parthenogenetic genus? Molecular Ecology 14: 2133-2142. PDF
Trewick SA, Goldberg J, Morgan-Richards M. 2005. Fewer species of Argosarchus and Clitarchus stick insects (Phasmida, Phasmatinae): evidence from nuclear and mitochondrial DNA sequence data. Zoologica Scripta 34: 483-491. PDF
Trewick SA, Morgan-Richard, M. 2005 After the deluge: mitochondrial DNA indicates Miocene radiation and Pliocene adaptation of tree and giant weta (Orthoptera: Anostostomatidae). Journal of Biogeography 32: 295-309. PDF
Trewick SA, Morgan-Richards M. 2005. New Zealand Wild: Stick Insects. Reed Publishing. Hong Kong.  
Trewick SA, Morgan-Richards M. 2004. Phylogenetics of New Zealand’s tree, giant and tusked weta (Orthoptera: Anostostomatidae): evidence from mitochondrial DNA. Journal of Orthoptera Research 13: 185-196. PDF
Trewick SA, Morgan-Richards M, Chapman HM. 2004. Chloroplast DNA diversity of Hieracium pilosella (Asteraceae) introduced to New Zealand: reticulation, hybridization and invasion. American Journal of Botany 91:73-85. PDF
Morgan-Richards M, Trewick SA, Chapman HM. 2004. Evidence of hybridisation among Hieracium species in New Zealand from flow cytometry.  Heredity 93 (1): 34-42. PDF
Morgan-Richards M, Wallis GP. 2003. Degree of cytogenetic differentiation fails to predict hybrid zone width in the weta  Hemideina thoracica (Orthoptera: Anostostomatidae). Evolution  57: 849-861. PDF
Morgan-Richards M. 2002. Fission or fusion? Mitochondial DNA phylogenetics of the chromosomal races of Hemideina crassidens (Orthoptera: Anostostomatidae). Cytogenetics & Genome Genetics 96: 217-222.  
Trewick SA, Morgan-Richards M, Henderson S, Pinter I, Russell SJ, Barrett JA, Gibby M, Vogel JC. 2002. Polyploidy, phylogeography and Pleistocene refugia of the rockfern Asplenium ceterach: evidence from cpDNA. Molecular Ecology 11: 2003-2012. PDF
Pinter I, Bakker F, Barrett J, Cox C, Gibby M, Henderson S, Morgan-Richards M, Rumsey F, Russell S, Trewick S, Schneider H, Vogel J. 2002. Phylogenetic and biosystematic relationships in four highly disjunct polyploid complexes in the subgenera Ceterach and Phyllitis in Asplenium (Aspleniaceae). Organisms, Diversity and Evolution 2: 299-311.  
Morgan-Richards M, King T, Trewick SA. 2001. The Evolutionary history of tree weta: a genetical approach. Chapter 7, Pp: 111-124. In: Field, L. [Ed] Biology of weta, King Crickets and their Allies. CABI Publishing, Oxford.  
Wallis GP, Trewick SA. 2001 Finding fault with vicariance: a critique of Heads (1998).  Systematic Biology 50:602-609. PDF
Morgan-Richards M, Gibbs GW. 2001. A phylogenetic analysis of New Zealand giant and tree weta (Orthoptera: Anostostomatidae: Deinacrida and Hemideina) using morphology and genetic characters. Invertebrate Taxonomy, 15: 1- 12. PDF
Morgan-Richards M, Trewick SA, Wallis GP. 2001. Chromosome races with Pliocene origins: evidence from mtDNA. Heredity, 86(3): 303-312. PDF
Trewick SA. 2001 Identity of an endangered grasshopper (Acrididae: Brachaspsis): taxonomy, molecules  and conservation. Conservation Genetics 2: 233-243. PDF
Trewick SA, Wallis GP. 2001. Bridging the ‘beech-gap’: invertebrate phylogeography implicates recent rather than ancient processes in New Zealand biogeographic patterns. Evolution 55:  2170-2180. PDF
Trewick SA. 2001. Scree weta phylogeography: surviving glaciation and implications for Pleistocene biogeography in New Zealand. New Zealand Journal of Zoology 28: 291-298. PDF
Steel DJ, Trewick SA, Wallis GP. 2000. Heteroplasmy of mitochondrial DNA in the Ophiuroid Astrobrachion constrictum. Journal of Heredity 91: 146-149. PDF
Trewick SA. 2000. Mitochondrial DNA sequences support allozyme evidence for cryptic radiation of New Zealand Peripatoides. Molecular Ecology 9: 269-282. PDF
Trewick SA. 2000. Molecular evidence for dispersal rather than vicariance as the source of flightless insects on the Chatham Islands, New Zealand. Journal of Biogeography 27: 1189-1200. PDF
Morgan-Richards M, Trewick SA, Wallis G. 2000. Characterization of a hybrid zone between two chromosomal races of the weta Hemideina thoracica following a geologically recent volcanic eruption. Heredity 85: 586–592. PDF
Trewick SA, Wallis, G.P. & Morgan-Richards, M. 2000.  Phylogeographic pattern correlates with Pliocene mountain-building in the alpine scree weta (Orthoptera, Anostostomatidae). Molecular Ecology 9: 657-666. PDF
Trewick SA, Morgan-Richards M. 2000. Artificial weta roosts: a technique for ecological study and population monitoring of tree weta (Hemideina) and other invertebrates. New Zealand Journal of Ecology, 24: 201-208. PDF
Trewick SA, Worthy T. 2000 Origins and prehistoric ecology of takahe: flightless Porphyrio (Aves: Rallidae). Chapter 4, Pp: 31-48. In: William G.L. and I.G. Jamieson (eds) The Takahe: 50 Years of Conservation Management and Research. Otago University Press, Dunedin.
Morgan-Richards M, Wolff K. 1999. Genetic structure and differentiation of Plantago major reveals a pair of sympatric sister species. Molecular Ecology, 8: 1027-1036. PDF
Trewick SA. 1999. A new weta from the Chatham Islands (Orthoptera: Raphidophoridae). Journal of the Royal Society of New Zealand 29: 165-173. PDF
Trewick SA. 1999. Molecular diversity of Dunedin peripatus (Onychophora: Peripatopsidae). New Zealand Journal Of Zoology 26: 381-393. PDF
Trewick SA. 1998. Kakapo: the paradoxical parrot. Nature Australia 26 (3): 54-63. PDF
Trewick SA. 1998. Sympatric cryptic species in New Zealand Onychophora. Biological Journal Linnean Society 63: 307-329. PDF
Morgan-Richards M. 1997. Intraspecific karyotype variation is not concordant with allozyme variation in the Auckland tree weta of New Zealand, Hemideina thoracica (Orthoptera: Stenopelmatidae). Biological Journal Linnean Society 60: 423-442.  
Trewick SA. 1997. Sympatric flightless rails Gallirallus dieffenbachii and G. modestus on the Chatham Islands, New Zealand; morphometrics and alternative evolutionary scenarios. J. Royal Soc. New Zealand, 27: 451-464. PDF
Trewick SA. 1997 Flightless and phylogeny amongst endemic rails (Aves: Rallidae) of the New Zealand region. Philosophical Transactions of the Royal Society, London B 352: 429-446. PDF
Trewick SA. 1997 On the skewed sex ratio of the kakapo Strigops habroptilus: Sexual and natural selection in opposition? Ibis 139:652-663. PDF
Trewick SA. 1996 Morphology and evolution of two takahe: Flightless rails of New Zealand. Journal of Zoology (London), 238: 221-237. PDF
Trewick SA, Morgan-Richards M. 1995 On the distribution of tree weta in the North Island, New Zealand. Journal Royal Society New Zealand 25: 485-493. PDF
Morgan-Richards M, Daugherty CH, Gibbs GW. 1995.  Specific status of the tree weta from Stephens Island, Mt Holdsworth and Mt Arthur, based on allozyme variation. Journal of the Royal Society of New Zealand, 25(2): 301-312. PDF
Morgan-Richards M. 1995. A new species of tree weta in the North Island of New Zealand (Orthoptera: Stenopelmatidae: Hemideina) New Zealand Entomologist, 18: 15-23. PDF
Morgan-Richards M, Townsend JA. 1995.  Hybridisation of tree weta on Banks Peninsula, New Zealand, and colour polymorphism within Hemideina ricta (Orthoptera: Stenopelmatidae). New Zealand Journal of Zoology, 22: 393-399. PDF
Trewick SA,  Dearden, P. 1994. A rapid protocol for DNA extraction and primer annealing for PCR sequencing. Biotechniques 17: 842-844. PDF
Trewick SA. 1990. Occurrence of Papilio, Graphium and Charaxes butterflies in Ugandan forests. Africa Journal of Ecology 28: 83-86. PDF
Trewick S, Wade PM. 1986. The distribution and dispersal of two alien species of Impatiens, waterway weeds in the Britisih Isles. Proceedings of the European Weed Research Society 7th Symposium on Aquatic Weeds 351–356. PDF